In a 78 yr-old forest stand in the woody part above ground per ha 286 kg N, 35 kg P, 154 kg K, 1115 kg Ca, 56 kg Mg and 48 kg S were accumulated. This refers to a phytomass of 141 ton dry matter. Shedding of leaves returns 59 kg N, 6.5 kg P, 22.4 kg K, 30.7 kg Ca, 7.4 kg Mg and 3.0 kg S into the soil. -from English summary